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To be able fully to appreciate trees, their mode of life, their enemies and their care, one must know something of their structure and life requirements.
Structure of trees: Among the lower forms of plants there is very little distinction between the various parts—no differentiation into root, stem, or crown. Often the lower forms of animal and vegetable life are so similar that one cannot discriminate between them. But as we ascend in the scale, the various plant forms become more and more complex until we reach the tree, which is the largest and highest form of all plants. The tree is a living organism composed of cells like any other living organism. It has many parts, every one of which has a definite purpose. The three principal parts are: the stem, the crown, and the root.
The stem: If we examine the cross-section of a tree, Fig. 86, we will notice that it is made up of numerous rings arranged in sections of different color and structure. The central part is known as the pith . Around the pith comes a dark, close-grained series of rings known as the heartwood , and outside the heartwood comes a lighter layer, the sapwood . The cambium layer surrounds the sapwood and the bark covers all. The cambium layer is the most important tissue of the tree and, together with part of the sapwood, transports the water and food of the tree. It is for this reason that a tree may be hollow, without heart and sapwood, and still produce foliage and fruit.
The crown: The crown varies in form in different species and is developed by the growth of new shoots from buds. The bud grows out to a certain length and forms the branch. Afterwards it thickens only and does not increase in length. New branches will then form from other buds on the same branch. This explains in part the characteristic branching of trees, Fig. 87.
The leaves are the stomach and lungs of the tree. Their broad blades are a device to catch the sunlight which is needed in the process of digesting the food of the tree. The leaves are arranged on the twigs in such a way as to catch the most sunlight. The leaves take up the carbonic acid gas from the air, decompose it under the influence of light and combine it with the minerals and water brought up by the roots from the soil. The resulting chemical combinations are the sugars and starches used, by the cambium layer in building up the body of the tree. A green pigment, chlorophyll , in the leaf is the medium by which, with the aid of sunlight, the sugars are manufactured.
The chlorophyll gives the leaf its green color, and this explains why a tree pales when it is in a dying condition or when its life processes are interfered with. The other colors of the leaf—the reds, browns and yellows of the fall or spring—are due to other pigments. These are angular crystals of different hues, which at certain times of the year become more conspicuous than at others, a phenomenon which explains the variation in the colors of the leaves during the different seasons.
It is evident that a tree is greatly dependent upon its leaves for the manufacture of food and one can, therefore, readily see why it is important to prevent destruction of the leaves by insects or through over-trimming.
The root: The root develops in much the same manner as the crown. Its depth and spread will vary with the species but will also depend somewhat upon the condition of the soil around it. A deep or a dry soil will tend to develop a deep root, while a shallow or moist soil will produce a shallow root, Fig. 88.
The numerous fine hairs which cover the roots serve the purpose of taking up food and water from the soil, while the heavy roots help to support the tree. The root-hairs are extremely tender, are easily dried out when exposed to the sun and wind, and are apt to become overheated when permitted to remain tightly packed for any length of time. These considerations are of practical importance in the planting of trees and in the application of fertilizers. It is these fine rootlets far away from the trunk of the tree that have to be fed, and all fertilizers must, therefore, be applied at points some distance from the trunk and not close to it, where merely the large, supporting roots are located. In the cultivation of trees the same principle holds true.
Requirements of trees: Trees are dependent upon certain soil and atmospheric conditions which influence their growth and development.
(1) Influence of moisture: The form of the tree and its growth and structure depend greatly upon the supply of moisture. Botanists have taken the moisture factor as the basis of classification and have subdivided trees into those that grow in moist places ( hydrophytes ), those that grow in medium soils ( mesophytes ), and those that grow in dry places ( xerophytes ). Water is taken up by the roots of the tree from the soil. The liquid absorbed by the roots carries in solution the mineral salts—the food of the tree—and no food can be taken up unless it is in solution. Much of the water is used by the tree and an enormous amount is given off in the process of evaporation.
These facts will explain some of the fundamental principles in the care of trees. To a tree growing on a city street or on a lawn where nature fails to supply the requisite amount of water, the latter must be supplied artificially, especially during the hot summer months, or else dead branches may result as seen in Fig. 89. Too much thinning out of the crown causes excessive evaporation, and too much cutting out in woodlands causes the soil to dry and the trees to suffer for the want of moisture. This also explains why it is essential, in wooded areas, to retain on the ground the fallen leaves. In decomposing and mixing with the soil, the fallen leaves not only supply the trees with food material, but also tend to conserve moisture in the ground and to prevent the drying out of the soil. Raking off the leaves from wooded areas, a practice common in parks and on private estates—hurts the trees seriously. Some soils may have plenty of moisture, but may also be so heavily saturated with acids or salts that the tree cannot utilize the moisture, and it suffers from drought just the same as if there had been no moisture at all in the soil. Such soils are said to be “physiologically dry” and need treatment.
In the development of disease, moisture is a contributing factor and, therefore, in cavities or underneath bandages where there is likely to be an accumulation of moisture, decay will do more damage than in places that are dry and exposed to the sun.
(2) Influence of soil: Soil is made up of fine particles of sand and rock and of vegetable matter called humus . A tree will require a certain soil, and unsuitable soils can be very often modified to suit the needs of the tree. A deep, moderately loose, sandy loam, however, which is sufficiently aerated and well supplied with water, will support almost any tree. Too much of any one constituent will make a soil unfit for the production of trees. If too much clay is present the soil becomes “stiff.” If too much vegetable matter is present, the soil becomes “sour.” The physical character of the soil is also important. By physical character is meant the porosity which results from breaking up the soil. This is accomplished by ploughing or cultivation. In nature, worms help to do this for the soil, but on streets an occasional digging up of the soil about the base of the tree is essential.
Humus or the organic matter in the soil is composed of litter, leaves and animal ingredients that have decayed under the influence of bacteria. The more vegetable matter in the humus, the darker the soil; and therefore a good soil such as one finds on the upper surface of a well-tilled farm has quite a dark color. When, however, a soil contains an unusual quantity of humus, it is known as “muck,” and when there is still more humus present we find peat . Neither of these two soils is suitable for proper tree growth.
(3) Influence of light: Light is required by the leaves in the process of assimilation. Cutting off some of the light from a tree affects its form. This is why trees grown in the open have wide-spreading crowns with branches starting near the ground as in Fig. 90, while the same species growing in the forest produces tall, lanky trees, free from branches to but a few feet from the top as in Fig. 91. Some trees can endure more shade than others, but all will grow in full light. This explains why trees like the beech, hemlock, sugar maple, spruce, holly and dogwood can grow in the shade, while the poplar, birch and willow require light. It also explains why, in the forest, the lower branches die and fall off—a process known in Forestry as “natural pruning,” The influence of light on the form of trees should be well understood by all those who plant trees and by those designing landscape effects.
(4) Influence of heat: Trees require a certain amount of heat. They receive it partly from the sun and partly from the soil. Evaporation prevents the overheating of the crown. The main stem of the tree is heated by water from the soil; therefore trees in the open begin growth in the spring earlier than trees in the forest because the soil in the open is warmer. Shrubs begin their growth earlier than trees because of the nearness of their crowns to their root systems. This also explains why a warm rain will start vegetation quickly. Too much heat will naturally cause excessive drying of the roots or excessive evaporation from the leaves and therefore more water is needed by the tree in summer than in winter.
(5) Influence of season and frost: The life processes of a tree are checked when the temperature sinks below a certain point. The tree is thus, during the winter, in a period of rest and only a few chemical changes take place which lead up to the starting of vegetation. In eastern United States, growth starts in April and ceases during the latter part of August or in early September. The different parts of a tree may freeze solid during the winter without injury, provided the tree is a native one. Exotic trees may suffer greatly from extreme cold. This is one of the main reasons why it is always advisable to plant native trees rather than those that are imported and have not yet been acclimatized. Frosts during mid-winter are not quite as injurious as early and late frosts and, therefore, if one is going to protect plants from the winter’s cold, it is well to apply the covering early enough and to keep it on late enough to overcome this difficulty.
The mechanical injuries from frost are also important. Snow and sleet will weigh down branches but rarely break them, while frost will cause them to become brittle and to break easily. Those who climb and prune trees should be especially cautious on frosty days.
(6) Influence of air: On the under side of leaves and on other surfaces of a tree little pores known as stomata may be found. In the bark of birch and cherry trees these openings are very conspicuous and are there known as lenticels . These pores are necessary for the breathing of the tree (respiration), whereby carbonic acid gas is taken in from the air and oxygen given out. The process of assimilation depends upon this breathing process and it is therefore evident that when the stomata are clogged as may occur where a tree is subjected to smoke or dust, the life processes of the tree will be interfered with. The same injurious effect results when the stomata of the roots are interfered with. Such interference may occur in cases where a heavy layer of soil is piled around the base of a tree, where the soil about the base of a tree is allowed to become compact, where a tree is planted too deep, or where the roots are submerged under water for any length of time. In any case the air cannot get to the roots and the tree suffers. Nature takes special cognizance of this important requirement in the case of cypress trees, which habitually grow under water. Here the trees are provided with special woody protuberances known as “cypress knees,” which emerge above water and take the necessary air. See Fig. 18.
Conclusions: From the foregoing it will be seen that trees have certain needs that nature or man must supply. These requirements differ with the different species, and in all work of planting and care as well as in the natural distribution of trees it is both interesting and necessary to observe these individual wants, to select species in accordance with local conditions and to care for trees in conformity with their natural needs.